Arrive in Puembo, a quiet rural neighbourhood of Quito located around 20 minutes from the airport and 45 minutes from the downtown city centre. Ecuador's mountain capital is, at 2,850m, the second highest capital in Latin America, lying in a narrow valley at the foot of the Pichincha Volcano. Arriving into the city by plane is quite spectacular as you approach over snow-capped mountains and extinct volcanoes - a window seat is recommended! You will be met on arrival at Mariscal Sucre International Airport (UIO) by a driver to take you to this evening's hotel, where more information will be given to you about tomorrow's schedule. The hotel is a tranquil, colonial-style property with a large garden and swimming pool. It's located in the countryside suburbs of Quito, near the airport; perfect for an early start tomorrow. The hotel can help organise a taxi for you into downtown Quito if you arrive early in the day and wish to visit the city.

Today we transfer to the airport early for our flight to the Galapagos Islands via Guayaquil (refueling stop only, you will not disembark the plane). Lying on the Equator, 970km west of the Ecuadorean coast, the Galapagos National Park is a unique collection of islands, formed by volcanoes over hundreds of thousands of years and consisting of six main islands, 12 smaller islands, and over 40 small islets, each one supporting a distinct and unique flora and fauna, including prehistoric creatures found nowhere else on earth. This desolate and fantastic habitat was discovered in 1535 by Fray Tomas de Berlanga. Charles Darwin visited the islands on the Beagle exactly three hundred years later, in 1835. He observed the total isolation of the giant reptiles and other creatures, and this played a substantial part in his theory of evolution. Of the extraordinary animals to be found in the Galapagos many are reptiles, such as the great tortoise, large land iguanas, numerous lizards and three species of non-poisonous snakes. There are also several species of turtle, which come ashore to mate. Up to three quarters of a million seabirds flock to the islands, including a third of the world's blue-footed boobies, frigatebirds, pelicans, cormorants, albatross and petrels. Only 35cm tall, the Galapagos penguin can be seen swimming among tropical mangroves, whilst offshore it is common to see dolphins, sea lions and fur seals playing in the water. Fearless though not tame, the extraordinary wildlife is often inquisitive and the opportunities for photography are superb. The Galapagos National Park charges a visitor fee of $100 USD (subject to change), payable in cash on arrival, which funds park maintenance and supervision in the Galapagos, as well as ecological study, conservation and infrastructure development in Ecuador's other national parks. Entry fees and the funds they generate for the national park system are among measures taken by the Ecuadorian government to protect its natural heritage. At San Cristobal Airport, a guide will meet us to help us collect our luggage. The easternmost island of the Galapagos archipelago and one of the oldest, San Cristobal's principal town is Puerto Baquerizo Moreno, the capital of the Galapagos. We'll make a visit to the interpretation centre where we will set the scene for the next week's trip, learning more about the Galapagos' history, geology, turbulent past and challenging future. A short, though often hot, uphill trail with steps, leads from the Interpretation Centre to the summit of Frigatebird Hill. Four panoramic viewpoints show Puerto Baquerizo Moreno from above, as well as the bays on both sides of the headland. On clear days even the distant, offshore sculpture of Kicker Rock can be seen. These wooded hilltops are one of the rare places where some magnificent and great frigatebirds form a mixed breeding colony. After our visit to the centre and Frigatebird Hill, we board our stylish vessel for the next week - the Archipel I.

After breakfast this morning we head out to explore Sante Fe island, giving us a chance to mingle with the iguanas. Located in the southeastern part of the Galapagos, this island is not volcanically formed, and is therefore mostly flat. Scientists theorise that this is the oldest island on the archipelago, and it is home to several endemic species including the Galapagos hawk and Galapagos mockingbird, as well as one of the two land iguana species on the islands. We'll disembark and explore on foot, where giant prickly pear cactus are dotted around. We continue onto South Plaza Island, one of the smallest islands in the Galapagos. A large colony of noisy sea lions, numbering about 1000 bulls, cows and pups, has its prime habitat on the smooth rocks. We will walk along a path through a cactus forest and view a combination of dry and coastal vegetation, populated by land iguanas, which can be seen sunning themselves or feeding on opuntia pads and fruits. Swallow-tailed gulls, which nest on the rugged southern cliffs, are usually seen, along with tropicbirds and Audubon's shearwaters. During the rainy season you can see the red sesuvium turn bright green and the leafless evening-blooming portulaca burst into large yellow flowers, which are loved by the iguanas.

Today we set sail for North Seymour Island, just north of Baltra Island, which is home to sea lions, marine iguanas, swallow-tailed gulls, magnificent frigatebirds and blue-footed boobies. North Seymour Island is probably the most exciting island photographically. Birdlife abounds, and close to the trail you will find many nesting pairs and young chicks. North Seymour is also home to the Galapagos' largest colony of frigatebirds. Their mating ritual is an ostentatious display; males expand the red sack at the base of their throat and perch atop a bush with wings fully extended, flapping furiously. Interested females circle overhead, and if so inclined, may join the male on terra firma. Further along the trail we can observe a colony of sea lions. In the afternoon the boat arrives to a small little island off the southern tip of Santiago Island called Sombrero Chino, or 'Chinese Hat', named for its unique shape. Chinese Hat comprises of a tilted cone and lava flows that run down to the coast. Here it is often possible to see Galapagos penguins and the abundant marine life, including rays, turtles and sea lions.

In the morning we arrive in Puerto Ayora, on Santa Cruz Island. Santa Cruz is the second largest island in the Galapagos and has the largest population, with Puerto Ayora as its main town. It also boasts the most varied of the islands' vegetation zones: coastal, transition, scalesia, miconia and pampa. The native scalesia forest of El Chato Tortoise Reserve is the best place to search for Galapagos giant tortoises in their most authentic setting. Though it can be quite moist and muddy, the visit will be an adventure, trying to spot the giant creatures silently leave their favourite ponds. While on the mainland, we'll also visit the Twin Craters - found on either side of the road leading to Puerto Ayora. These impressive formations were formed by magma domes, which hardened on the outside while the lava continued flowing inside, leaving huge and empty magma chambers that eventually collapsed, leaving two large holes. The craters lie within a lush scalesia cloud forest, a high-altitude plant species that are endemic to the Galapagos. This area is also home to the carpenter finch, which uses tools to search for food. This afternoon we visit the Fausto Llerena Breeding Centre, where we can observe tortoises that come from various islands of the archipelago - at Fausto Llerena they are breeding tortoises only to repopulate areas where their populations have been affected by non-native wildlife (predominantly feral dogs). One of the main attractions here is the embalmed Lonesome George - estimated to have died at between 93 and 109 years of age, Lonesome George is recognised as the last of its kind, and as a symbol of the efforts invested into conserving endangered species.

We land on Isabela Island today, and walk to Moreno Point this morning. Located on the north coast between the volcanos of Sierra Negra and Cerro Azul, the trail runs along a solidified lava flow into a complex of coastal lagoons, where we'll find an abundance of birdlife around the lakes and mangroves. In the afternoon we'll Visit Urbina Bay, located at the base of Alcedo Volcano. Hike the trail to get a view of the caldera. Walk among a massive marine reef that was raised out of the water in 1950 during an uplift, and see sculptures of dried coral and other ocean formations while touring the area on foot. Urbina Bay also offers great opportunities to see land iguanas that have astonishingly yellow skin, flightless cormorants, land tortoises as well as marine turtles and rays.

This morning's excursion takes us to Punta Espinoza on Fernandina, the youngest of the Galapagos Islands. Witness the large colony of marine iguanas and a variety of bird life, including the unique flightless cormorant. This bird had to adjust its way of survival and perfect its skills of finding food in the ocean. Their wings, tails and feet progressively adapted for swimming. To see these birds is to witness evolution happening right in front of you. We'll tour a number of different trails and follow the path along the beach and across lava flows, for a unique opportunity to explore one of the least-visited areas of the Galapagos. While having lunch we sail to Vicente Roca point back on Isabela Island, just at the mouth of Isabela's seahorse-shape. From small panga boats we'll enter a dark cave below a spectacular arch, accompanied by roaring echoes of the waves. Just around the corner the collapsed amphitheatre of Volcan Ecuador offers another impressive view. The calmer waters of the coves are well-protected against the ocean swell and are fairly cold, but a great place for snorkelling amongst various species of shark, penguins, puffer fish and seahorses.

In the morning, we will travel to the black-sand beach of Egas Port on Santiago Island to witness the striking and fascinating giant lava formations. Very few plants have managed to survive on this island due to its harsh environment and relatively new lava flow. Take an afternoon excursion to the red sand beaches of Rabida to see a lagoon and the famous bachelor sea lion colony. This area is known for its birdlife, including pelicans, white-cheeked pintails and boobies. Explore the trails of Rabida; a short path leads to a saltwater lagoon; another trail goes past the lagoon to the interior, where the revered palo santo trees grow. When burned, the branches of this tree give off a pleasing aroma and ward off mosquitoes. Grab a snorkel and discover many species of fish and other marine life. Keep an eye out for white-tip sharks and sea lions.

Explore the mangrove lagoon at Black Turtle Cove by a small panga. In the afternoon, we will transfer to the airport for our flight to Quito. On arrival back on the mainland, you'll have a transfer to the hotel for your final night in Ecuador.

The trip ends after breakfast at the hotel in Puembo, Quito. There are no activities planned today, so you are free to depart at any time. Pick-up times for your airport transfer to Mariscal Sucre International Airport (UIO) will be re-confirmed locally. If your flight is departing later in the day, luggage storage facilities are available at the hotel. If you would like to book additional nights in Quito at the end of your stay, these can be arranged either at the hotel in Puembo, or in a hotel in downtown Quito. We recommend spending extra time in Quito, or adding to your nature-based holiday by booking our four-night Amazon extension package (trip code: AE).